Navegando por Assunto "Edessinae"
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Item Acesso aberto (Open Access) Análise cladística de Edessa (Hypoxys) com a descrição de um grupo novo de espécies (Heteroptera, Pentatomidae, Edessinae)(Universidade Federal do Pará, 2018-12-10) MENDONÇA, Maria Thayane da Silva; FERNANDES, José Antônio Marin; http://lattes.cnpq.br/6743352818723245Genus Edessa has many problems of taxonomy and nomenclature, but to make the revision of the genus in only one study becomes impracticable due to the large number of species (<600 species), so it was proposed to review small groups of species. The subgenus Hypoxys is a monophyletic group according to the single phylogenetic analysis, supported by four synapomorphies. This subgenus of Edessa has 17 described species and more than 40 new species for science, according to an unpublished revision. Thus, this work aims to propose a group of species within Hypoxys, to describe 15 new species, and to make an identification key for them, as well as to run a cladistic analysis to Hypoxys. Thereby, we intend to test the monophyly of Hypoxys and relationship among species with the inclusion of more species and new characters. Thirty-five specimens from national and foreign institutions and private collections were analyzed. These specimens were described, measured and photographed. For the cladistic analysis, a data matrix with 35 taxa and 39 characters was made using Mesquite software, the matrix was calculated in TNT and the cladogram was edited using WinClada. The maximum parsimony was used to run the cladistic analysis. To reduce the final number of trees implicit weighing (k = 2 to k = 6) was used. To verify the values of clade support, the relative bremer support and the symmetric resampling method were calculated. The analysis without implicit weighing resulted in 73 trees with 78 steps, consistency index 57 and retention index 89. The analysis with implicit weighing (k = 2 to k = 6) resulted in 37 trees with 76 steps, consistency index 59 and retention index 90. Tree topology was similar using or not implicit weighing. All resulting trees from relative bremer and symmetric resampling showed high branch support for Hypoxys and the target group. The results corroborate Hypoxys as monophyletic, being formed by three large groups, which share seven synapomorphies and three homoplasic synapomorphies: ventral coloration of the humeral angles of color; scutellum with dark spot in the anterior third present; apex of the dorsal rim fused to the margin of the posterolateral angles; excavation in rounded metapleuron; no roughness of the textured surface of the evaporatorium; presence of the dorsal rim with a thickness variation from the center to the posterolateral angles; format of the straight paramere; presence of a flap in the ventral region of the proctiger; presence of a "carpet of hairs" on the ventral rim; expansion of the ventral rim of the tumid. In addition, the target group has other additional diagnostic features, they are: the posterior region of the hyaline pronotum, brown, extending to the posterior margin of the pronotum; conexivum without stains; two pairs of circular, dark spots on the ventral part of the thoracic surrounded by a greenish ring; ventral region evenly green with a central yellowish color in the abdomen; dark and circular spots next to the pseudosutures; posterior margin of the gonocoxite 8 projected later. The result of the analysis showed the clade Hypoxys as monophyletic, as well as the target group of this work composed by 15 new species.Item Acesso aberto (Open Access) Análise cladística e descrição de um novo gênero de Edessinae (Hemiptera, Heteroptera e Pentatomidae)(Universidade Federal do Pará, 2016-04-06) NUNES, Benedito Mendes; FERNANDES, José Antônio Marin; http://lattes.cnpq.br/6743352818723245Item Acesso aberto (Open Access) Análise cladística e revisão do subgênero nominal de Edessa (Heteroptera: Pentatomidae: Edessinae)(Universidade Federal do Pará, 2017-01-23) SILVA, Valéria Juliete da; FERNANDES, José Antônio Marin; http://lattes.cnpq.br/6743352818723245Pentatomidae is the fourth numerous and diverse family of Heteroptera. From Pentatomidae, Edessinae has more than 300 described species. Edessinae is a Neotropical subfamily with nine genera: Edessa Fabricius, 1803, Brachystethus Laporte, 1832, Ascra Say, 1832, Peromatus Amyot & Serville, 1843, Olbia Stål, 1862, Pantochlora Stål, 1870, Doesburgedessa Fernandes, 2010, Paraedessa Silva & Fernandes, 2013 and Grammedessa Correia & Fernandes, 2016, genera with well-defined diagnoses except Edessa which is considered a deposit of species to the subfamily. Edessa has the greatest taxonomical and nomenclatural problems of Edessinae, because it historically has a great confusion with the limits of the genus and subfamily. To solve the problem was proposed the review of Edessa from groups of species and subgenera. Edessa is composed by five subgenera: Aceratodes Amyot & Serville, 1843, Dorypleura Amyot & Serville, 1843, Pygoda Amyot & Serville, 1843, Hypoxys Amyot & Serville, 1843and the nominal subgenus. Among subgenera only the nominal was not reviewed. As Edessa (Edessa) is an “empty taxa”, composed only by the type specie, it were used the characters of the groups of species by Stål (1872) as start point to the delimitation of this subgenus. During the bibliography research was found a mistake in the typification of Edessa, so here the type specie was changed from E. cervus (Fabricius, 1787) to E. antilope (Fabricius, 1798). To recognize and limit Edessa and to create relationship hypothesis among the species was realized a cladistics analysis. It were included the species considered by Stål (loc. cit.) as part of the Edessa group of species, as well as species that possess the characters mentioned by Stål as diagnosis to this group. The data matrix is composed for 111 morphological characters and 85 taxa, 13 of outgroup and 72 of ingroup. For the analysis were realized two weighting schemes: equal and implicit weight of characters with K varying from 3–12, and two types of search: traditional and new technologies. The cladogram with K=8 and traditional search have 763 steps, IC: 19 and IR: 60. Based on this cladogram the subgenus Edessa is composed by 10 species known for the science: E. antilope, E. cervus, E. taurina Stål, 1862, E. ibex Breddin, 1903, E. arabs (Linnaues, 1758), E. cylindricornis Stål, 1872, E. rondoniensis Fernandes & van Doesburg, 2000, E. burmeisteri Fernandes & van Doesburg, 2000, E. cerastes Breddin, 1905 and E. elaphus Breddin, 1905, and six new morphotypes: E. sp. nov. “near flavinernis”, E. sp. nov. “close flavinernis”, E. sp. nov. “near 112”, E. sp. nov. “close 112”, E. sp. nov. “40” and E. sp. nov. “131”. The subgenus Edessa is diagnosed by the mainly green color on dorsal surface; humeral angles at least twice longer than large, black apex of humeral angle posteriorly curved, whole or bifid; embolium contrasting in color to the corium; corium with at least one yellow vein; dorsal rim of pygophore narrow and continuous with the base of posterolateral angles. Also the cladistics analysis suggests the monophyly of Edessinae and polyphyly of Edessa in its current composition. Changes in taxonomical status to genera of Aceratodes, Dorypleura, Pygoda and Hypoxys are corroborated. Peromatus appears with an internal branch in the analysis; it reinforces the necessity to review the genus. Also were recognized and described 13 new group of species to Edessa. Were redescribed species known to the science and described new species to the genus. Nomenclatural problems have been identified, with 11 synonyms proposed and a revalidation of taxon previously in synonymy; lectotypes were designated and a key of identification of species is given.Item Acesso aberto (Open Access) Caracterização do grupo Edessa rufomarginata e descrição de sete novas espécies (Heteroptera, Pentatomidae, Edessinae)(2006-09) SILVA, Eduardo José Ely e; FERNANDES, José Antônio Marin; GRAZIA, JocéliaThe Edessa group rufomarginata is proposed to include E. rufomarginata (De Geer, 1773), E. marginalis (Dallas, 1851), E. albomarginata (Stål, 1855), E. abdominalis Erichson, 1848, E. corallipes Erichson, 1848, E. aulacosterna Stål, 1872, E. ovalis Stål, 1872, and E. nigropunctata Berg, 1884. Seven new species are described in this group, six from Brazil and one from Argentina: E. brasiliensis sp. nov., E. castaneolineata sp. nov., E. cerradensis sp. nov., E. chapadensis sp. nov., E. luteovenulata sp. nov., E. rufodorsata sp. nov. (Argentina) and E. virididorsata sp. nov. These species are known from Venezuela, Suriname, Guyana, Brazil, Paraguay and Argentina.Item Acesso aberto (Open Access) Descrição de um gênero novo e três espécies novas de Edessinae (Hemiptera: Heteroptera: Pentatomidae)(Universidade Federal do Pará, 2017-12-07) ALMEIDA, Flávio Roberto de Albuquerque; FERNANDES, José Antônio Marin; http://lattes.cnpq.br/6743352818723245The family Pentatomidae is the fourth most diverse among Heteroptera, comprising about 4.700 species divided in, approximately, 800 genera. The family is a monophyletic taxa characterized by the loss of gonapophyses 8, gonapophyses 9 reduced and fused to gonocoxites 9, gonangulum absent and the ductus receptaculi dilated and invaginated, forming three distinct walls. The subfamily Edessinae is the second most diverse, comprising about 320 species divided in nine genera: Edessa Fabricius, 1803 (about 300 species), Ascra Say, 1832 (14 species), Brachystethus Laporte, 1832 (10 species), Peromatus Amyot & Serville, 1843 (seven species), Olbia Stål, 1862 (three species), Pantochlora Stål, 1870 (one species), Doesburgedessa Fernandes, 2010 (five species), Paraedessa Silva & Fernandes, 2013 (nine species) and Grammedessa Correia & Fernandes, 2016 (12 species). The genus Edessa has many taxonomic and nomenclatural problems related to the lack of a reliable diagnosis. In this way, species which have general characteristics of the subfamily and did not fit in the diagnostic features of other genera of Edessinae are considered to belonging to Edessa. This practice made the genus a "deposit" of species with indefinite limits. The proposal for the genus is to revise Edessa in groups of species sharing morphological distinctive characters (possible synapomorphies). One of these enigmatic groups of Edessa is formed by Edessa celsa Distant, 1890 and three allied new species. Here we describe a new genus for these four species sharing a unique set of characteristics and compare with Hypoxys, subgenus of Edessa with similar facies. Diagnosis, illustrations of the metasternal process and external and internal genitalia, photographs of the dorsal and ventral view, and a distributional map are provided. A key to the genera belonging to Edessinae and a key to the species of Plagaedessa are also provided. Male and female genitalia of Plagaedessa celsa n. comb. are described for the first time. Distribution of P. celsa is extended to Costa Rica and Mexico. The lectotype of Edessa celsa is here designated.Item Acesso aberto (Open Access) Proposição de um novo gênero de Edessinae (Heteroptera, Pentatomidae)(Universidade Federal do Pará, 2015-02-12) CORREIA, André Oliveira; FERNANDES, José Antônio Marin; http://lattes.cnpq.br/6743352818723245Edessinae have seven genera, among them Edessa is the largest with more than 250 names proposed and more than 300 species to be described. The diversity resulted in a complex taxonomical history, so Edessa is the genus with more taxonomical and nomenclatural problems in Edessinae. Review the genus in groups of species is the best proposal to organize Edessa and it already resulted in description of two new genera of Edessinae. This study selected three species described by Beddin (1905), in addition to eight new species chosen for morphological characters. A new genus to Edessinae is proposed, uniting species with four black and punctured stripes longitudinal on dorsal surface of the head; humeral angles laterally projected, slightly flattened dorso-ventrally; metasternal process with branches of bifurcation narrow, long, thin in the apex; abdominal trichobothria placed laterally, not aligned with spiracles; gonocoxites 8 and laterotergites 8 always punctured in females; gonocoxites 8 large, subequal in length to laterotergites 9; pygophore with dorsal rim with semicircular excavation, paramere posteriorly projected, genital cup superior process dark, laminar, partially fused to the cup process. Edessa pallicornis, E. bugabensis, E. rorativentris and E. stillativentris are redescribed and transferred to the new genus. E. strigiceps is proposed as junior synonym of Gen. nov. pallicornis. This study gives the first description of males of Gen. nov. bugabensis and Gen. nov. rorativentris. Extended distribution of Gen. nov. bugabensis (Costa Rica), Gen. nov. rorativentris (Brazil), Gen. nov. stillativentris (Colombia). The lectotype of Gen. nov. bugabensis is here designated. Eight new species are described to the new genus: Gen. nov. sp. n. 203a, Gen. nov. sp. n. 203e, Gen. nov. sp. n. 205, Gen. nov. sp. n. 206, Gen. nov. sp. n. 207, Gen. nov. sp. n. 207c, Gen. nov. sp. n. 297 e Gen. nov. sp. n. 297b.Item Acesso aberto (Open Access) Urhai gen. nov., um novo gênero para Edessinae (Hemiptera: Heteroptera: Pentatomidae)(Universidade Federal do Pará, 2025-03) FERREIRA, Adiney Ferreira; FERNANDES, José Antônio Marin; http://lattes.cnpq.br/6743352818723245; https://orcid.org/0000-0001-7450-5296Edessinae is the second most diverse subfamily of Pentatomidae, with approximately 500 described species distributed in 17 genera. This taxon is considered monophyletic, supported by molecular evidence and morphological characteristics. Among the genera, Edessa stands out for harboring more than 300 species, but it faces taxonomic challenges, including problems of nomenclature and classification, and is often considered a "species repository". This work aims to describe a new genus with three species previously allocated to Edessa and five new species. For the study, 94 specimens from national and foreign institutions were analyzed. Urhai gen. nov. is easily recognized by the morphology of the body, mainly presenting the head with median punctuations forming two vertical stripes on the clypeus, antennal articles I–III with small dark circular spots, pronotum with developed humeral angle; flattened and punctuated dorsoventrally; distal margin laminar; posterior angle spiniform, posterior margin of the pronotum covering the anterior part of the scutellum, metasternal process flattened; anterior arms of the bifurcation reaching or nearly reaching the limit of the propleura; distal margin generally straight. Diagnoses, species identification key and distribution maps were presented. This study is of great importance in understanding the diversity of Edessinae.